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The Peregrine Falcon (Falco peregrinus), also known simply as the Peregrine,Heinzel, H., Fitter, R., & Parslow, J. (1995). Birds of Britain and Europe, 5th ed. Collins ISBN 0-00-219894-0. and historically also as "Duck Hawk" in North America, is a cosmopolitan bird of prey in the family Falconidae. It is a large falcon, about the size of a large crow, with the female being larger than the male,Snow et al. (1998) and with a blue-gray back, barred white underside, and a black head and "mustache". About seventeen subspecies are recognized, which vary in appearance and range.
The breeding range includes land regions from the Arctic tundra to the Tropics. Essentially, this species can be found everywhere on Earth, except in the polar regions, on very high mountains, in deserts, and most tropical rainforests making it the world's most widespread falcon, and in fact the most widespread bird of prey.J. Ferguson-Lees, D. A. Christie: Raptors of the World. Christopher Helm, London 2001. ISBN 0-7136-8026-1 The only major ice-free landmass from where it is entirely absent is New Zealand. Both the English and scientific names of this species mean "wandering falcon" and refer to the migratory habits of some populations of this widespread species.
It feeds almost exclusively on medium-sized birds, but will occasionally hunt small mammals. It reaches sexual maturity at one year, and mates for life. It nests in a scrape, normally on cliff edges or, in recent times on tall man-made structures.Cade, T. J., M. Martell, P. Redig, T. Septon & H. Tordoff: Peregrine Falcons in urban North America. In: D. M. Bird, D. E. Varland & J. J. Negro (eds.): Raptors in Human Landscapes. London, San Diego; 1996: S. 313 The Peregrine Falcon became an endangered species due to the use of pesticides, especially DDT. Since the ban on DDT from the beginning of the 1970s onwards, the populations recovered, supported by large scale protection of nesting places and releases to the wild.T. J. Cade, J. H. Enderson, C. G. Thelander & C. M. White (Eds): Peregrine Falcon Populations Their management and recovery. The Peregrine Fund, Boise, Idaho, 1988. ISBN 0-9619839-0-6
The Peregrine Falcon has a body length of 3450 cm and a wingspan of around 80120 cm . The male and female have similar markings and plumage, but as in many birds of prey the Peregrine Falcon displays marked sexual dimorphism in size, with the female measuring up to 30 percent larger than the male. Males weigh 440750 g, and the noticeably larger females weigh 9101500 g; for variation in weight between subspecies, see under that section below.
The back and the long, pointed wings of the adult are usually bluish black to slate grey with indistinct darker barring (see "Subspecies" below); the wingtips are black. The underparts are white to rusty and barred with thin clean bands of dark brown or black. The tail, colored like the back but with thin clean bars, is long, narrow and rounded at the end with a black tip and a white band at the very end. The top of the head and a "mustache" along the cheeks are black, contrasting sharply with the pale sides of the neck and white throat. The cere is yellow, as are the feet, and the beak and claws are black. The upper beak is notched near the tip, an adaptation which enables falcons to kill prey by severing the spinal column at the neck. The immature bird is much browner with streaked, rather than barred, underparts, and has a pale bluish cere.
Taxonomy and systematics
This species was first described by Marmaduke Tunstall in his 1771 Ornithologia Britannica under its current binomial name. The scientific name Falco peregrinus, means "wandering falcon" in Latin. Indeed, the species' common name refers to its wide-ranging flights in most European languages.E.g. French faucon , German falke, Italian falco , Polish sokol wedrowny, Slovak sokol stahovavy, Swedish pilgrimsfalk The Latin term for falcon, falco, is related to falx, the Latin word meaning sickle, in reference to the silhouette of the falcon's long, pointed wings in flight.
The Peregrine Falcon belongs to a genus whose lineage includes the hierofalconsContra Helbig et al. (1994), Wink et al. (1998). The supposed basal position of the hierofalcons was due to them having a cytochrome b numt: see Wink & Sauer-Gurth (2000) and the Prairie Falcon . This lineage probably diverged from other falcons towards the end of the Late Miocene or in the Early Pliocene, about 85 million years ago (mya). As the Peregrine-hierofalcon group includes both Old World and North American species, it is likely that the lineage originated in western Eurasia or Africa. Its relationship to other falcons is not clear; the issue is complicated by widespread hybridization confounding mtDNA sequence analyses; for example a genetic lineage of the Saker Falcon is knownHelbig et al. (1994), Wink et al. (1998) which originated from a male Saker producing fertile young with a female Peregrine ancestor some 100,000 years ago.[ref]
Today, Peregrines are regularly hybridized in captivity with other species such as the Lanner Falcon to produce the "perilanner", a somewhat popular bird in falconry as it combines the Peregrine's hunting skill with the Lanner's hardiness, or the Gyrfalcon to produce large, strikingly-colored birds for the use of falconers. As can be seen, the Peregrine is still genetically close to the hierofalcons, though their lineages diverged in the Late Pliocene .Helbig et al. (1994), Wink et al. (1998), Griffiths (1999), Wink & Sauer-Gurth (2000), Groombridge et al. (2002), Griffiths et al. (2004), Nittinger et al. (2005)
Numerous subspecies of the Peregrine Falcon have been described, with 19 accepted by the Handbook of the Birds of the World.Vaurie (1961), White (1994), Snow et al. (1998)
The nominate subspecies Falco peregrinus peregrinus, described by Tunstall in 1771, breeds over much of temperate Eurasia between the tundra in the north and the Pyrenees, Mediterranean region and Alpide belt in the south. It is mainly non-migratory in Europe, but migratory in Scandinavia and Asia. Males weigh 580750 g, while females weigh 9251,300 g. It includes brevirostris, germanicus, rhenanus, and riphaeus. Falco peregrinus calidus, described by Latham in 1790, was formerly called leucogenys and includes caeruleiceps. It breeds in the Arctic tundra of Eurasia, from Murmansk Oblast to roughly Yana and Indigirka Rivers, Siberia. It is completely migratory, and travels south in winter as far as sub-Saharan Africa. It is paler than peregrinus, especially on the crown. Males weigh 588740 g, while females weigh 9251,333 g. Falco peregrinus japonensis, described by Gmelin in 1788, includes kleinschmidti and pleskei, and harterti seems to refer to intergrades with calidus. It is found from northeast Siberia to Kamchatka (though it is possibly replaced by pealei on coast there), and Japan. Northern populations are migratory, while those of Japan are resident. It is similar to peregrinus, but the young are even darker than those of anatum.
Falco peregrinus macropus, described by Swainson in 1837 is the Australian Peregrine Falcon. It is found in Australia in all regions except the southwest. It is non-migratory. It is similar to brookei in appearance, but is slightly smaller and the ear region is entirely black. The feet are proportionally large. Falco peregrinus submelanogenys described by Mathews in 1912, is the Southwest Australian Peregrine Falcon. It is found in southwest Australia and is non-migratory.
Falco peregrinus peregrinator, described by Sundevall in 1837, is known as the Indian Peregrine Falcon, Black Shaheen, or Indian Shaheen.The shaheen of Arabic and Persian writers are usually Barbary Falcons; those in Indian and Pakistani sources normally refer to peregrinator. It was formerly sometimes known as Falco atriceps or Falco shaheen. Its range includes South Asia from Pakistan across India to Sri Lanka and Southeastern China. It is non-migratory. It is small and dark, with rufous underparts barred with lighter color. In Sri Lanka this species is found to favour the higher hills while the migrant calidus is more often seen along the coast.
Falco peregrinus anatum, described by Bonaparte in 1838, is known as the American Peregrine Falcon, or "Duck Hawk"; its scientific name means "Duck Peregrine Falcon". At one time, it was partly included in leucogenys. It is mainly found in the Rocky Mountains today. It was formerly common throughout North America between the tundra and northern Mexico, where current reintroduction efforts seek to restore the population. Most mature anatum, except those that breed in more northern areas, winter in their breeding range. Most vagrants that reach western Europe seem to belong to the more northern and strongly migratory tundrius, only considered distinct since 1968. It is similar to peregrinus but is slightly smaller; adults are somewhat paler and less patterned below, but juveniles are darker and more patterned below. Males weigh 500570 g, while females weigh 900960 g.Falco peregrinus cassini, described by Sharpe in 1873, is also known as the Austral Peregrine Falcon. It includes kreyenborgi, the Pallid FalconAlso called "Kleinschmidt's Falcon", but this might equally refer to F. p. kleinschmidti which is a junior synonym of japonensis, a leucistic morph occurring in southernmost South America, which was long believed to be a distinct species. Its range includes South America from Ecuador through Bolivia, northern Argentina and Chile to Tierra del Fuego and Falkland Islands. It is non-migratory. It is similar to nominate, but slightly smaller with a black ear region. The variation kreyenborgi is medium grey above, has little barring below, and has a head pattern like the Saker Falcon, but the ear region is white.
Falco peregrinus pealei, described by Ridgway in 1873, is also known as Peale's Falcon, and includes rudolfi. It is found in the Pacific Northwest of North America, northwards from the Puget Sound along the British Columbia coast (including the Queen Charlotte Islands), along the Gulf of Alaska and the Aleutian Islands to the far eastern Bering Sea coast of Russia. It is possibly found on the Kuril Islands and the coasts of Kamchatka as well. It is non-migratory. It is the largest subspecies, and it looks like an oversized and darker tundrius or like a strongly barred and large anatum. The bill is very wide. Juveniles occasionally have pale crowns. Falco peregrinus tundrius, described by C.M. White in 1968, was at one time included in leucogenys It is found in the Arctic tundra of North America to Greenland. It migrates to wintering grounds in Central and South America. Most vagrants that reach western Europe belong to this subspecies, which was previously united with anatum. It is the New World equivalent to calidus. It is smaller than anatum. It is also paler than anatum; most have a conspicuous white forehead and white in ear region, but the crown and "moustache" are very dark, unlike in calidus. Juveniles are browner, and less grey, than in calidus, and paler, sometimes almost sandy, than in anatum.
Falco peregrinus madens, described by Ripley and Watson in 1963, is unusual in having some sexual dichromatism. If the Barbary Falcon (see below) is considered a distinct species, it is sometimes placed therein. It is found in the Cape Verde Islands, and is non-migratory; it is endangered with only six to eight pairs surviving. Males have a rufous wash on crown, nape, ears and back; underside conspicuously washed pinkish-brown. Females are tinged rich brown overall, especially on the crown and nape. Falco peregrinus minor was first described by Bonaparte in 1850. It was formerly often perconfusus. It is sparsely and patchily distributed throughout much of sub-Saharan Africa and widespread in Southern Africa. It apparently reaches north along the Atlantic coast as far as Morocco. It is non-migratory, and small and dark. Falco peregrinus radama, described by Hartlaub in 1861, is found in Madagascar and Comoros. It is non-migratory.
Falco peregrinus brookei, described by Sharpe in 1873, is also known as the Mediterranean Peregrine Falcon or the Maltese Falcon. It includes caucasicus and most specimens of the proposed race punicus, though others may be pelegrinoides, Barbary Falcons (see also below), or perhaps the rare hybrids between these two which might occur around Algeria. They occur from the Iberian Peninsula around the Mediterranean, except in arid regions, to the Caucasus. They are non-migratory. It is smaller than the nominate subspecies, and the underside usually has rusty hue. Males weigh around 445 g, while females weigh up to 920 g.
Falco peregrinus ernesti, described by Sharpe in 1894, is found from Indonesia to Philippines and south to Papua New Guinea and Bismarck Archipelago. Its geographical separation from nesiotes requires confirmation. It is non-migratory. It differs from the nominate in the very dark, dense barring on its underside and its black ear coverts. Falco peregrinus furuitii, described by Momiyama in 1927, is found on the Izu and Ogasawara Islands. It is non-migratory. It is very rare, and may only remain on a single island. It is a dark form, resembling pealei in color, but darker, especially on tail. Falco peregrinus nesiotes described by Mayr in 1941, is found in Fiji and probably also Vanuatu and New Caledonia. It is non-migratory.
Falco peregrinus pelegrinoides, first described by Temminck in 1829, is found in the Canary Islands through north Africa and the Near East to Mesopotamia. It is most similar to brookei, but is markedly paler above, with a rusty neck, and is a light buff with reduced barring below. It is smaller than the nominate subspecies; females weigh around 610 g. Falco peregrinus babylonicus described by P.L. Sclater in 1861, is found in eastern Iran along the Hindu Kush and Tian Shan to Mongolian Altai ranges. It is paler than pelegrinoides, and somewhat similar to a small, pale Lanner Falcon. It is smaller than Peregrine Falcon; males weigh 330400 g, while females weigh 513765 g. These last two races are often split as Barbary Falcon Falco pelegrinoides. There is a 0.60.7% genetic distance in the Peregine-Barbary Falcon ("peregrinoid") complex.Wink et al. (2000) These birds inhabit arid regions from the Canary Islands along the rim of the Sahara through the Middle East to Central Asia and Mongolia. They have a red neck patch but otherwise differ in appearance from the Peregrine proper merely according to Gloger's Rule. The Barbary Falcon has a peculiar way of flying, beating only the outer part of its wings like fulmars sometimes do; this also occurs in the Peregrine, but less often and far less pronounced.Snow et al. 1998. The Barbary Falcon's shoulder and pelvis bones are stout by comparison with the Peregrine, and its feet are smaller. They have no postzygotic reproduction barriers in place, but they breed at different times of year than neighboring Peregrine Falcon subspecies.Helbig et al. (1994)Snow et al. (1998)Wink et al. (1998)Wink et al. (2000)Wink et al. (2004)
Ecology and behavior
The Peregrine Falcon lives mostly along mountain ranges, river valleys, coastlines, and increasingly in cities. In mild-winter regions, it is usually a permanent resident, and some individuals, especially adult males, will remain on the breeding territory. Only Populations that breed in arctic climes typically migrate great distances during the northern winter.
The Peregrine Falcon is often stated to be the fastest animal on the planet in its hunting dive, the stoop, which involves soaring to a great height and then diving steeply at speeds of over hitting one wing of its prey, so as not to harm itself on impact. A study testing the flight physics of an 'ideal falcon' found a theoretical speed limit at 400 km/h for low altitude flight and 625 km/h for high altitude flight. Despite these theoretical values, measurements of real stoops by using radar resulted in maximum diving speeds of only .T. Alerstam: Radar observations of the stoop of the Peregrine Falcon Falco peregrinus and the Goshawk Accipiter gentilis. Ibis 169, 1986: S. 267273
The life span in the wild is up to 15.5 years. Mortality in the first year is between 5970%, declining to between 2532% in adults. Apart from anthropogenic threats like collision with man-made objects, the Peregrine may be killed by large eagles or large Owls.
The Peregrine Falcon is host to a range of parasites and pathogens. It is a vector for Avipoxvirus, Newcastle disease virus, Falconid herpesvirus 1 (and possibly other Herpesviridae), and some mycoses and bacterial infections. Endoparasites include Plasmodium relictum (usually not causing malaria in the Peregrine Falcon), Strigeidae trematodes, Serratospiculum amaculata (nematode), and tapeworms. Known Peregrine Falcon ectoparasites are chewing liceColpocephalum falconii which was described from specimens found on the Peregrine Falcon, Colpocephalum subzerafae, Colpocephalum zerafae and Nosopon lucidum (all Menoponidae), Degeeriella rufa (Philopteridae), Laemobothrion tinnunculi (Laemobothriidae). All are known from other Falco species too. Ceratophyllus garei (a flea), and Hippoboscidae flies .Raidal et al. (1999), Raidal & Jaensch (2000), Dewey & Potter (2002), Dalgleish (2003)
The Peregrine Falcon feeds almost exclusively on medium sized birds such as doves, waterfowl and songbirds. Other than bats taken at night, it rarely hunts small mammals, but will on occasion take rats, voles, hares, mice and squirrels; the coastal populations of the large subspecies pealei feed almost exclusively on seabirds. In the Brazilian mangrove swamp of Cubatao, a wintering falcon of the subspecies tundrius was observed while hunting successfully a juvenile Scarlet Ibis Insects and reptiles make up a small proportion of the diet, which varies greatly depending on what prey is available. In urban areas, the main item of the Peregrine's diet is the feral pigeon, followed by other common city birds such as Common Starlings and Common Swifts.
The Peregrine Falcon hunts at dawn and dusk, when prey are most active, but in cities also nocturnally, particularly during migration periods when hunting at night may become prevalent. Nocturnal migrants taken by Peregrines include species as diverse as Yellow-billed Cuckoo, Black-necked Grebe, Virginia Rail and Common Quail. It requires open space in order to hunt, and therefore often hunts over open water, marshes, valleys, fields and tundra. It searches for prey either from a high perch or from the air.Ehrlich et al. (1992) Once prey is spotted, it begins its stoop, folding back the tail and wings, with feet tucked. The air pressure from a dive could possibly damage a bird's lungs, but small bony tubercles in a falcon's nostrils guide the shock waves of the air entering the nostrils (compare intake ramps and inlet cones of jet engines), enabling the bird to breathe more easily while diving by reducing the change in air pressure. Prey is struck and captured in mid-air; the Peregrine Falcon strikes its prey with a clenched foot, stunning or killing it, then turns to catch it in mid-air. The Peregrine will drop it to the ground and eat it there if it is too heavy to carry. Prey is plucked before consumption.
The Peregrine Falcon is sexually mature at the end of the first year of age but in healthy populations they breed after two to three years of age. The pair mates for life and returns to the same nesting spot annually. The courtship flight includes a mix of aerial acrobatics, precise spirals, and steep dives. The male passes prey it has caught to the female in mid-air. To make this possible, the female actually flies upside-down to receive the food from the male's talons. The Peregrine Falcon is territorial during the breeding season; nesting pairs are usually more than 1 km apart, and often much farther, even in areas with large numbers of pairs. The distance between nests ensures sufficient food supply for pairs and their chicks. Within a breeding territory, a pair may have several nesting ledges; the number used by a pair can vary from one or two to seven in a 16 year period. The pair defends the chosen nest site against other Peregrines, and often against eagles or ravens.
The Peregrine Falcon nests in a scrape, normally on cliff edges or, today regularly in many parts of its range, on tall buildings or bridges. Cliff nests are generally located under an overhang, on ledges with vegetation, and south-facing sites are favored. In some regions, as in parts of Australia and on the west coast of Northern North-America, large tree hollows are used for nesting. Before the demise of most European peregrines, there was a large population of peregrines in central and western Europe using the disused nests of other large birds. The female chooses a nest site, where she scrapes a shallow hollow in the loose soil, sand, gravel, or dead vegetation in which to lay eggs. No nest materials are added. In remote, undisturbed areas such as the Arctic, steep slopes and even low rocks and mounds may be used as nest sites. The man-made structures used for breeding closely resemble the natural cliff ledges that the Peregrine prefers for its nesting locations.
Mostly three to four eggs (range 1-5) are laid in the scrape. The eggs are white to buff with red or brown markings. They are incubated for 29 to 33 days, mainly by the female. The male also helps with the incubation of the eggs over day, but at night only the female incubates. The date of egg-laying varies according to locality, but is generally from February to March in the Northern Hemisphere, and from July to August in the Southern Hemisphere (the Australian subspecies macropus may breed as late as November and equatorial populations may nest anytime between June and December). The female generally lays another clutch if the eggs are lost early in the nesting season, though this is extremely rare in the Arctic owing to the short summer season. As a result of some infertile eggs and natural losses of nestlings, the average number of young found in nests is 2.5, and the average number that fledges is about 1.5.
After hatching, chicks are covered with creamy-white down and have disproportionately large feet. The male, which is called the "tiercel", brings food to the female and chicks, but the chicks are fed by the female, which stays at the nest and watches the young. The hunting territory of the parents can extend a radius of 19 to 24 km (12-15 miles) from the nest site. Chicks fledge 42 to 46 days after hatching, and remain dependent on their parents for up to two months.
Relationship with humans
See also: Peregrine Falcons in popular culture
The Peregrine Falcon became an endangered species because of the use of pesticides, especially DDT during the 1950s, 60s, and 70s.T. J. Cade, J. H. Enderson, C. G. Thelander & C. M. White (Eds): Peregrine Falcon Populations Their management and recovery. The Peregrine Fund, Boise, Idaho, 1988. ISBN 0-9619839-0-6 Pesticide biomagnification interfered with reproduction, thinning eggshells and reducing the number of eggs that survived to hatching. The organochlorine build-up in the falcon's fat tissues would result in less calcium in the eggshells, leading to flimsier, more fragile eggs. In several parts of the world, such as the eastern USA and Belgium, this species became extinct as a result. Peregrine eggs and chicks are often targeted by black marketeers and unscrupulous egg collectors, so it is normal practice not to publicize unprotected nest locations.
The Peregrine Falcon was used in falconry for more than 3,000 years, beginning with nomads in central Asia. Due to its ability to dive at high speeds, it was highly sought-after and generally used by experienced falconers. Peregrine Falcons are also occasionally used to scare away birds at airports to reduce the risk of bird-plane strikes, improving air-traffic safety. The Peregrine Falcon is featured on the Idaho State Quarter, in recognition of its efforts in its recovery.
In the USA, Canada and Germany, Wildlife services in Peregrine Falcon recovery teams breed the species in captivity. The chicks are usually fed through a chute or with a hand puppet mimicking a Peregrine's head, so they cannot see to imprint on the human trainers. Then, when they are old enough, the rearing box is opened, allowing the bird to train its wings. As the fledgling gets stronger, feeding is reduced forcing the bird to learn to hunt. This procedure is called hacking back to the wild. To release a captive-bred falcon, the bird is placed in a special cage at the top of a tower or cliff ledge for some days or so, allowing it to acclimate itself to its future environment.
Worldwide recovery efforts have been remarkably successful. The widespread restriction of DDT use eventually allowed released birds to breed successfully. The Peregrine Falcon was removed from the U.S. Endangered Species list on August 25, 1999.
Many Peregrine Falcons have settled in large cities, including London, Ontario, Derby, Brisbane and Cologne, and all across the U.S., where they nest on cathedrals, skyscraper window ledges, and the towers of suspension bridges. At least 18 pairs nested in New York City proper in 2005. In Virginia, state officials working with students from the Center for Conservation Biology of the College of William and Mary in Williamsburg successfully established nesting boxes high atop the George P. Coleman Memorial Bridge on the York River, the Benjamin Harrison Memorial Bridge and Varina-Enon Bridge on the James River, and at other similar locations. Thirteen new chicks were hatched in this Virginia program during a recent year. Over 250 falcons have been released through the Virginia program. In the UK, there has been a recovery of populations since the crash of the 1960s. This has been greatly assisted by conservation and protection work led by the RSPB. Peregrines now breed in many mountainous and coastal areas, especially in the west and north, and nest in some urban areas, capitalizing on the urban pigeon populations for food.
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(1994): 60. Peregine Falcon. In: (editors): Handbook of Birds of the World, Volume 2 (New World Vultures to Guineafowl): 274275, plate 28. Lynx Edicions, Barcelona. ISBN 84-87334-15-6
(1994): Family Falconidae. In: (editors): Handbook of Birds of the World, Volume 2 (New World Vultures to Guineafowl): 216275, plates 2428. Lynx Edicions, Barcelona. ISBN 84-87334-15-6
(2000): Advances in the molecular systematics of African raptors. In: : Raptors at Risk: 135147. WWGBP/Hancock House, Berlin/Blaine. PDF fulltext
(1998): Molecular systematics of holarctic raptors (Order Falconiformes). In: : Holarctic Birds of Prey: 2948. Adenex & WWGBP. PDF fulltext
(2000): Phylogenetic relationships between Black Shaheen (Falco peregrinus peregrinator), Red-naped Shaheen (F. pelegrinoides babylonicus) and Peregrines (F. peregrinus). In: : Raptors at Risk: 853857. WWGBP/Hancock House, Berlin/Blaine. PDF fulltext
(2004): Phylogenetic relationships in the Hierofalco complex . In: : Raptors Worldwide: 499504. WWGBP, Berlin. PDF fulltext
(2003): Peregrine Falcon (Falco peregrinus). Version of 2003-01-07. Retrieved 2007-08-13.
Falcon Research Group
Peregrine Falcon FundThe Canadian Peregrine Foundation
Video and other media of Peregrines:
Peregrine Falcon videos on the Internet Bird Collection
The only Peregrine Falcon Webcam in the Southern Hemisphere
The Raptor Resource Project. Peregrine falcon, eagle, owl, and kestrel cams.
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This article is licensed under the GNU Free Documentation License. It uses material from the Wikipedia article Peregrine Falcon